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...ician.<ref>{{cite book|editor=Steed, H. E.|title=The register of Tonbridge School from 1826 to 1910|year=1911|publisher=Rivingtons|url=https://archive.org/de ...{{sfn|Starr|2003|p=4}} and was educated at [[Tonbridge School]] (1870–75). At [[Trinity College, Cambridge]], he graduated B.A. as 4th [[Wrangler (Univer ...

...e field of [[biomechanics]]. For thirty years he was Professor of Zoology at the [[University of Leeds]]. ...ton.lib.cam.ac.uk/vwebv/holdingsInfo?bibId=30333 }}</ref> His PhD research at [[University of Cambridge|Cambridge]] was supervised by [[James Gray (zoolo ...